Xenorophus |
We can put some limits on this. At the younger end, since all odontocetes echolocate, it's unlikely to have evolved any later than their last common ancestor, which is estimated to have lived around 34 million years ago. On the other hand, it's notable that the toothless, baleen, whales do not use ultrasound; in fact, they are specialised in the exact opposite direction to produce sounds well below, not above, the range of human hearing. This suggests that they evolved along different lines, and that the origins of ultrasonic echolocation lie somewhere after the two split.