Antilopine Antelopes: Dik-diks

Kirk's dik-dik

The "antilopine" antelopes are, as their name might suggest, mostly typical examples of their kind. I suspect that when many people think of antelopes in general, gazelles and springbok are among the first to come to mind (although, to be fair, so might say, impalas). Traditionally, the antilopines have been divided into two main groups: the gazelle-like antelopes and the dwarf antelopes, with the latter surely being the less familiar to those of us living outside of Africa. Both groups have at least some species that don't fit the "typical" image of the subfamily. For the gazelle-like animals, that's probably the gerenuk and its relative, the dibatag.

Among the dwarf antelopes, we have the dik-diks.

While some researchers subdivide them further, four species of dik-dik are widely recognised, at least three of which are reasonably common within their respective homelands. This may partly be because they are too small to be worth hunting for meat - although, inevitably. this still happens from time to time. 

Antilopine Antelopes: Dwarf Antelopes of Eastern and Central Africa

Oribi

The term "dwarf antelope" can reasonably be applied to many different kinds of antelope that are smaller than, say a typical goat. In this common sense, it's not a precise term, and could refer to species that belong to quite different branches of the antelope family tree. When mammalian zoologists used the term in the 20th century, however, it was often more precise, referring to those thought to be most closely related to the gazelles as a branch within the "antilopine" subfamily of "typical" antelopes.

Technically, these dwarf antelopes were collectively referred to as "neotragines" and assumed to be a natural grouping within the antelopes more widely. Genetic analysis over the last couple of decades has muddied these waters considerably, not least because the genus for which the branch was named, Neotragus, turns out not to be closely related to the gazelles, and is something else entirely. Even if we look solely at those dwarf antelopes that we can still say belong to the antilopine subfamily, it turns out that they don't have a single common ancestor. Specifically, one of them is more closely related to gazelles than it is to any of the other members of its purported evolutionary branch. 

Antilopine Antelopes: The Largest Gazelles

Grant's gazelle

The word "gazelle", as used in everyday English is a little vague, referring to a general concept of slim, agile, antelopes but not to anything with a precise scientific definition. Strictly speaking, however, it refers to a specific group of closely related animals, and some species that are commonly called "gazelles" strictly speaking aren't. During the 20th century, all true gazelles were placed in a single genus, Gazella, and it's this that defines the more rigid definition of what does and doesn't count. 

In more recent decades, Gazella has been split in three, with the resurrection of two 19th-century names as "new" genera. The original Gazella is widespread, including animals from both Asia and North Africa, but the other two are exclusively African. Eudorcas (literally "true antelope" in Ancient Greek) includes Thomson's gazelle of the Serengeti and its various relatives, while the remaining genus is Nanger, whose name apparently comes from a local Senegalese word for one of the species.

Antilopine Antelopes: The Gazelles of Asia

Arabian gazelles

I suspect that, on the whole, westerners associate gazelles with Africa. We think of the ones we see in wildlife documentaries, being pursued by cheetahs or leopards across the plains of the Serengeti or similar places. However, the most current theory suggests that they may have originated in Asia and various species survive on both continents today, having split apart around 2 or 3 million years ago at or shortly before, the start of the Ice Ages.

How many species that might be is still a matter of debate, and much of it centres on what's probably the first part of Asia you'd think of to look for desert-dwelling animals: the Middle East. For much of the 20th century, there were generally regarded as being two species living here, not counting one or two then thought to be extinct. And then, well, all that fell apart for reasons I wrote about on this blog back in 2013.

Antilopine Antelopes: Gazelles of North Africa

Dorcas gazelle

(Brief note: My internet connection was down for three days over the weekend, which is why this post is delayed from the usual.)

One of the things that most distinguishes gazelles from other kinds of antelope is that they are adapted to dry environments. They don't come much drier than the Sahara so it should be little surprise that gazelles are relatively common here. In fact, the dorcas gazelle (Gazella dorcas) is one of the most widespread of all gazelle species, being found right across the Sahara from the Atlantic to the Red Sea, as well as further south along the Red Sea coast in Eritrea and Djibouti and across the Sinai into extreme southern Israel. In the north, it's largely restricted to the eastern parts of the Mediterranean coast, being absent from northern Algeria, Morocco, and Tunisia.

When the Desert is Too Dry

The round-tailed ground squirrel lives
further east, and is not threatened

Many mammal species are territorial, carving out a patch of land for themselves which they then defend from same-sex members of their own species. Typically, they are less bothered about members of the opposite sex, for obvious reasons, and such territories will often overlap. Male territories tend to be larger than those defended by females, making it easier for them to meet as many females as possible. 

The size and relative location of such territories naturally vary between species, but also depend on the local conditions of terrain, climate and so on. The harder it is to find food, for instance, the larger your territory will need to be. As young animals grow up and leave home, they will need to find unoccupied territories to inhabit, or else somehow drive an existing resident out and take over. Males commonly travel further than females so that they don't end up with only their sisters or close cousins as potential mating partners, although there are a few species where it works the opposite way around.

Bats in the Fynbos

Lesser horseshoe bat

From a human perspective, the ability of bats to navigate in pitch darkness through the use of biosonar is undoubtedly remarkable. Not only do they have to be able to avoid obstacles, but also, in the case of the insect-eating species, to locate and identify tiny prey items against the background. Their ability to do this is affected by a number of environmental factors, and the exact nature of the echolocation calls, and how the bat interprets them, may affect which of those factors have the most influence.

For example, the distance that a biosonar ping can travel is affected by both the temperature and humidity of the air through which it travels. Since there are bats in almost every habitat suitable for insects to live, we might expect the way that they use their sonar to vary depending on the local climate.

Small Cats: The Domestic Cat's Closest Relatives

Sand cat

The range of species included in the genus Felis has changed significantly over the years. Because of the way such things are named, it includes, by definition, the domestic cat and all its closest relatives - but how close is close?

Unsurprisingly, the genus appears in the very first catalogue of scientific names, all the way back in 1758. At the time, it included every one of the seven species of cat known to the author - wild/domestic cats, lions, tigers, leopards, jaguars, lynxes, and ocelots. Over the next few decades, whenever a new cat species was described (as you might imagine, cheetahs and pumas were among the first) it was added to the same genus. At the time, the modern concept of "taxonomic family" didn't exist, and, round about the same time that families became a thing and the Felidae were named, the big cats were hived off into a genus of their own, Panthera.

The Dog Family: Foxes of Africa

Fennec fox

The fact that foxes will eat pretty well anything that's small enough has meant that some species have been able to colonise surprisingly harsh environments. The kit fox, for example, inhabits the desert shrublands of much of the western US, while Blanford's fox lives in the dry hills of the Middle East and south-central Asia. No fox, however, is more desert adapted than the fennec fox (Vulpes zerda) of North Africa.

Fennec foxes (sometimes simply called "fennecs") are also among the most distinctive of foxes. For one thing, they're the smallest wild members of the dog family, with particularly small adults as little as 33 cm (13 inches) in length, plus tail, and weighing just 800 g (28 oz.)  They have pale sandy-and-white fur, which is unusually long and soft - they even have fur on the pads of their feet, to give them some protection from baking hot sand. And, of course, there are the huge ears, quite out of proportion to the rest of the animal, which help to radiate away heat in an animal that would rather not lose too much water by panting.

Fennec foxes live across almost the whole of the Sahara Desert, from the Atlantic coast to the Nile valley. They are not typically found east of the Nile, where the closely related Blanford's foxes are found instead, but there are a few exceptions, and, for example, both species inhabit the Sinai. In fact, fennec foxes actually prefer the open sand dunes of the desert interior, an exceptionally harsh and arid environment.