Oligocene (Pt 8): The First Tapir and the Last Hoofless Horse

Miohippus

A close look at the evolutionary history of horses reveals that it's more complicated than sometimes presented, with numerous side branches forming a bushy tree of different species, many of which ultimately died out without descendants. This applies both to the origins of the group in its early days and its period of great diversification through the Miocene and Pliocene epochs. In the Oligocene, however, the picture, at least so far as we can tell, was rather simpler.

We know of two genera of horse that made it into the Oligocene from the preceding epoch. Mesohippus had been around for a while, Miohippus was a relative newcomer, appearing towards the very end of the Eocene. The two are rather similar, to the point that it has been argued they should be treated as different species of the same genus (which would be Miohippus, as that was named first) and they are both found in fossil beds across the United States and southern Canada, with Miohippus being known from Washington state to Florida and Mesohippus primarily in the west. The similarity also led to proposals early on that the one directly evolved into the other, but it's now clear that they lived alongside one another for millions of years, which scuppers that idea.

Oligocene (Pt 4): Time of the Giants

Paraceratherium

The Grande Coupure saw many new animals pouring across from Asia into Europe. Whether or not the tapirs were among them is not immediately obvious. The oldest tapir that we know of is a fossil discovered in Germany and dated to the early Oligocene, not long after the Coupure. Named Protapirus, we know that it, or its very close relatives were found not only in Asia, but also North America, by the second half of the epoch and managed to survive, with relatively little change, into the Early Miocene that followed, dying out about 20 million years ago. 

At first glance, since the oldest fossil is German, it appears that tapirs originated in Europe and then spread east, and it's purely a coincidence that they happened to do so after the Coupure - which, after all, was a time of climatic change. The problem is, there wasn't anything remotely tapir-like living in Europe before the Coupure, but there were plenty of potential ancestors elsewhere. So it's perhaps more likely that the first true tapirs were Asian, and we simply haven't found their fossils yet. Even so, we can at least say that Protapirus, and its later relative Paratapirus (which never seems to have left Europe) really were tapirs, rather than some close relative. A key feature here is that, unlike their earlier relatives, they already had the modifications to the nasal bones that suggest the presence of the short trunk that modern tapirs have, although it was probably less prominent than in current species.

The Jungles of British Columbia

It's changed a bit since...

Earth's climate has changed dramatically over the billions of years of its existence. The recent warming events may be unusually fast, but they are by no means unusually large. Not much more than 12,000 years ago the world was in the grip of the Ice Ages, when the polar ice caps stretched much further than today, and places like southern Italy and northern California were cold, sparsely forested steppeland.

On the other hand, there were times when the world was much hotter than today, warm enough that there appear to have been no ice caps at all, even at the poles. During the Age of Mammals - the eon of time since the extinction of the non-avian dinosaurs - the hottest of these was the Eocene Climactic Optimum. This took place a little over 50 million years ago, and it lasted rather a long time, punctuated by short periods of even greater heat. (That's 'short' as in they lasted tens of thousands of years, but not millions).