Oligocene (Pt 7): Not Quite Camels, Not Quite Pigs

Protoceras

While the ruminants of Oligocene North America would have looked similar to the musk deer of today, some of the other cloven-hoofed mammals inhabiting the continent at the time were more distinctive. Protoceratids no longer survive, but they had already been around for millions of years at the dawn of the Oligocene, and would survive throughout the whole of the following epoch and a little way into the one after that - an impressive record. Despite this, they never seem to have been very common, and the only undoubted Oligocene example is Protoceras, known primarily from Nebraska, Wyoming, and South Dakota.

It remains unclear exactly what protoceratids were, beyond the fact that they were obviously related to other cloven-hoofed animals. Some features suggest that they were closely related to ruminants (as was assumed when they were first discovered in the 19th century) while others indicate a close relationship to camels; it may even be that they are some early branch that doesn't fit well with either. Despite being the animal for which the group is named, Protoceras is not so well known as its later relatives, many of which notable for possessing a third horn on their snouts in addition to those in the place we'd expect to find horns on a goat or antelope. 

Miocene (Pt 15): The Coming of the Rain Shadow

Ramoceros

As the world passed into the Late Miocene, worldwide temperatures dipped from the high summer of the Middle Miocene towards... well, if not autumn, perhaps a late summer, since the world was still overall, probably warmer than it is today. For North America, however, perhaps a more significant effect was caused by the slow shuffling of the continents. A large chunk of rock that corresponds, broadly speaking, to present-day California began to tilt, causing the eastern edge to rise up.

We're not talking a huge tilt, of the sort that you'd notice if you were standing on it - it's been estimated that the eastern end was rising by somewhere around 0.25 mm (1/1000 of an inch) per year relative to the western one. Give it a few million years though, and that starts to be noticeable. While this process really only accelerated during the subsequent, Pliocene, epoch, even at this time, the rise of the eastern end of that block of crust was slowly, but surely, creating what we now call the Sierra Nevada Mountains.

Miocene (Pt 10): The Beasts with Three Horns

Syndyoceras

At the dawn of the Miocene, 23 million years ago, North America was, in many respects, quite different from the way it is today. It was, at the time, separated from South America by a wide expanse of sea, and, perhaps more significantly, the Sierra Nevada and Cascade Mountain ranges, which today run from southern British Columbia down into Mexico, did not yet exist as any more than a series of particularly high hills. While the Rockies were already mountainous, so that there was at least some rain shadow, in general, the continent was wetter and more forested than today, lacking the great grassy prairies that are so dominant today.

As the climate warmed, sea levels rose, flooding low-lying coastal regions and turning major valleys into bays. For much of the epoch, for instance, Florida was almost entirely underwater. Further north, however, it was still connected to Asia (and Greenland, although that's less significant in the grand scheme of things). Perhaps because the land bridge was both far to the north and relatively mountainous, not many animals seem to have crossed it, but that changed in the Early Miocene as the climate began to improve, and only increased as the epoch wore on. And, for whatever reason, most of them were heading east - into America.