Oligocene (Pt 7): Not Quite Camels, Not Quite Pigs

Protoceras

While the ruminants of Oligocene North America would have looked similar to the musk deer of today, some of the other cloven-hoofed mammals inhabiting the continent at the time were more distinctive. Protoceratids no longer survive, but they had already been around for millions of years at the dawn of the Oligocene, and would survive throughout the whole of the following epoch and a little way into the one after that - an impressive record. Despite this, they never seem to have been very common, and the only undoubted Oligocene example is Protoceras, known primarily from Nebraska, Wyoming, and South Dakota.

It remains unclear exactly what protoceratids were, beyond the fact that they were obviously related to other cloven-hoofed animals. Some features suggest that they were closely related to ruminants (as was assumed when they were first discovered in the 19th century) while others indicate a close relationship to camels; it may even be that they are some early branch that doesn't fit well with either. Despite being the animal for which the group is named, Protoceras is not so well known as its later relatives, many of which notable for possessing a third horn on their snouts in addition to those in the place we'd expect to find horns on a goat or antelope. 

Oligocene (Pt 3): From Musk Deer to Hell-Pigs

Entelodon

The Grande Coupure that marked the beginning of the Oligocene in Europe did not have as dramatic an effect on the cloven-hoofed animals as it did on some other mammalian groups. But the change was nonetheless noticeable, with several primitive forms that had once inhabited the continent suddenly dying out as it merged with Asia. Others, however, survived through the break, and the absence of their earlier competitors may even have helped them prosper.

These include the gelocids, which first appeared close to the end of the previous epoch. Few of the known fossils of these animals are in good condition, and there is some debate as to whether they are a true group of animals at all, or just a vague term used to collect similar-looking creatures that we can't place elsewhere. That aside, we can at least say that they physically resembled (but were probably not related to) musk deer. That is, they were relatively small, hornless animals with long legs suited for running fast, but lacking the large fang/tusks that mark true musk deer. They did well enough that some, such as Pseudogelocus, are known not only from France and Germany, but also Mongolia, suggesting that they crossed over in the opposite direction to most other mammal groups.

Prehistoric Mammal Discoveries 2022

New reconstruction of the sabretooth cat
Homotherium, showing that the teeth
would not have been as visible as popularly
supposed

2022 approaches its conclusion and I have to say that, while far from perfect, it was an improvement on the previous two years. There were a couple of weeks without posts this year, one for positive reasons, and one less so, but overall it has been good. Next year, you can look forward to something other than monkeys in my "family" series of posts and also to the conclusion of the long-running Miocene series, which should be as early as February. Until then, here is the now-traditional year-end look back at the paleontological discoveries that didn't quite make it into the blog.

Large Herbivores

Probably the most distinctive thing about deer is that the males have antlers; branching bony head ornaments that are shed and regrown each year. This naturally raises the question of how this evolved, since no other animal has quite the same thing. Acteocemas was an Early Miocene deer, but despite living very early in the group's evolutionary history, it already had antlers that split into two near the tip - which the horns of animals such as cows and true antelopes never do. A Spanish fossil of the antlers described earlier this year showed that it was already shed and regrown, but microscopic analysis indicated that it appeared to have been present for over a year, suggesting longer a more irregular pattern of shedding that must have changed to the seasonal pattern we are familiar with more recently, perhaps in the Middle Miocene.

Miocene (Pt 11): Horses on the Grasslands

Daeodon

The lush greenery of Early Miocene North America was a good place for large mammalian herbivores. Many of these, such as musk deer, pronghorns and camels, were, in one fashion or another, cud-chewing animals, able to extract maximum nutrition from a grassy or leafy diet. But many, of course, were not, either finding different ways to get the most out of their food, or else going for plants that were generally easier to digest.

Some of these were, like the ruminants, cloven-hoofed animals. Today, the main group of non-ruminant cloven-hoofed animals are the pigs, but they have never truly lived wild in the Americas, with feral 'razorbacks' only having arrived with the white man. Instead, America has peccaries, also known as javelinas, animals that look very much like pigs, but have a number of crucial differences.